Interesting article. I completely agree that animals can grieve. We recently lost our oldest cat, Peepers. He was 19 and was having alot of problems (not eating, throwing up, etc); we had to put him to sleep. I was very difficult for both my husband and I; both of us cried over our loss. Our other 2 cats were very needy for the next few days, always wanting to be petted, sitting beside us and so on. They also took to sleeping in Peepers favorite spots. This lasted about 2 weeks. I really do think they were missing him.
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